These measurements do have an association with whole carcass composition. Williams et al. (2018a, 2018b) demonstrated the capacity for either GR tissue depth combined with lamb hot carcass weight (figure 4.6.1a) or rib fat depth combined with beef hot carcass weight (figure 4.6.1b) to predict the percentage of fat in the carcass. In both cases there were strong associations, but there was also significant error.